Crinoid List and Photos

Crinoids are echinoderms that have been around at least since the Early Ordovician and are abundant in the ocean today. Modern crinoids are stalkless (comatulids) in shallow water and stalked in deep (several hundred meters+). Paleozoic crinoids were shallow-water and only rarely stalkless.

Complete specimens from holdfast to crown are rare but well-documented. Crinoid calices (sgl. calyx) are best described as “locally common.” That is to say, they’re rare in general, but where they are found, they are often abundant. This is because they are associated with mass mortalities – where storms bury an area quickly smothering everything. Crinoids with large bodies may be crushed.

I’ve been involved with three papers that named ten new crinoid species. Details and photos will be posted eventually.

The most common fossils from crinoids are the columns or individual rings (called “Indian beads”) called columnals. Body plates are common, but usually unidentifiable. Dr. Tom Kammer researched crinoid body plates in the Button Mold Knob Member of the New Providence Shale and was able to match most of them to known species, plus a new species or two, as well.

Preservation Key: C = Calcified, Ph = Phosphatic, P = Pyritized, S = Silicified, 1= w/o, matrix, 2 = w/ matrix

# Not available for sale, just for identification

CRINOIDS

Abrotocrinus debrae Ausich, Goldstein & Yates# – named for my wife in a paper published in 2000. Second known specimen – destined for the Smithsonian to join the first. Unidentifiable before being prepped, this specimen survived a fire at the prep lab and smelled like smoke when it was returned to me! Muldraugh Fm., Middle Mississippian, Hardin Co., Kentucky.

 Abrotocrinus debrae - I'm a co-author of this new species.
Abrotocrinus debrae – I’m a co-author of this new species.
Plaster cast of holotype of  Abrotocrinus debrae. Original at the U.S. National Museum (Smithsonian).
Plaster cast of holotype of Abrotocrinus debrae. Original at the U.S. National Museum (Smithsonian).

Acrocrinus shumardi Yandell# – goblet-shaped body, Indian Springs Shale, Upper Mississippian, Crawford Co., IN – C,2

 Acrocrinus shumardi with Archimedes bryozoan
Acrocrinus shumardi with Archimedes bryozoan

Actinocrinites jugosus (Hall)# – is a very large crinoid with ornate plates. Muldraugh Fm., Middle Mississippian, Hardin Co., Kentucky. Prepped by Chris Wright.

Actinocrinites jugosus  is a very large crinoid with ornate plates. Muldraugh Fm., Middle Mississippian, Hardin Co., Kentucky. Prepped by Chris Wright.
Actinocrinites jugosus (Hall)

Agassizocrinus conicus (Shumard) – found as a basal cone of a stalkless crinoid. Upper Mississippian, Grayson Co., KY – C,1,2

Agassizocrinus conicus (Shumard) - found as a basal cone of a stalkless crinoid. Upper Mississippian
Agassizocrinus conicus (Shumard) cluster of basal cones

Alisocrinus carleyi (Hall) = Glyptocrinus or Dimerocrinus? – Waldron Shale, Middle Silurian, Clark Co., Indiana

Alisocrinus carleyi - patterned calyx plates
Alisocrinus carleyi – patterned calyx plates

Ancyrocrinus bulbosus (Hall) – Beechwood Limestone, Givetian, Middle Devonian, Clark Co., IN – S,1

Ancyrocrinus bulbosus - side view
Ancyrocrinus bulbosus – side view
Ancyrocrinus bulbosus - top view
Ancyrocrinus bulbosus – top view

Aorocrinus cassedayi (Lyon)# – Beechwood Limestone, Givetian, Middle Devonian, Clark Co., IN – S,1

Aorocrinus cassedayi - basal cup
Aorocrinus cassedayi – basal cup

Bicidiocrinus wetherbyi (Wachsmuth & Springer) – spines abundant in the Glen Dean Fm., Upper Mississippian, Grayson Co., KY – C,1

Bicidiocrinus wetherbyi spines
Bicidiocrinus wetherbyi spines

Barycrinus cornutus (Owen & Shumard) – New Providence Shale, Middle Mississippian, KY – C,1

 Barycrinus cornutus body plates
Barycrinus cornutus body plates

Barycrinus sculptilis Hall – have ornamented plates, New Providence Shale, Middle Mississippian, KY – C,1

Barycrinus sculptilis - an assortment of body plates
Barycrinus sculptilis – an assortment of body plates

Batocrinus crassitestus Rowley# – is found in the Salem Limestone and the Somerset Shale Mbr., Middle Mississippian. This specimen is from Hardin Co., Kentucky. C,1

Batocrinus crassitestus, dampened to show detail, 3 cm view in microscope.
Batocrinus crassitestus, dampened to show detail, 3 cm view in microscope.

Botryocrinus polyxo (Hall) – crinoid cup, Waldron Shale, Middle Silurian, Clark Co., Indiana, C,1,2

Botryocrinus polyxo - has a distinctive cup in the Waldron Shale
Botryocrinus polyxo – has a distinctive cup in the Waldron Shale
Botrocrinus polyxo in Waldron Shale
Botrocrinus polyxo

Callicrinus longispinus Weller# – a mold from the Laurel Dolostone, Bardstown, Nelson Co., Kentucky. Found by randomly cracking a small boulder.

 Callicrinus longispinus - the calcite plates dissolved away.
Callicrinus longispinus – the calcite plates dissolved away.

Catillocrinus tennesseae Shumard# – has odd-shaped body plates, New Providence Shale, Middle Mississippian, KY – C,P,1

 Catillocrinus tennesseae body plates
Catillocrinus tennesseae body plates

Cosmetocrinus gracilis Kirk# – Muldraugh Fm., Middle Mississippian, Hardin Co., Kentucky – C,2

Cosmetocrinus gracilis among Platycrinites georgii
Cosmetocrinus gracilis among Platycrinites georgii

Cyathocrinites astralus Kammer – New Providence Shale, Middle Mississippian, KY – C,P,1

Cyathocrinites astralus Kammer - New Providence Shale, Middle Mississippian
Cyathocrinites astralus body plates

Cyathocrinites iowensis (Owen & Shumard)# and Synbathocrinus swallovi Hall, Muldraugh Fm., Middle Mississippian, Hardin Co., Kentucky

Cyathocrinites iowensis and Synbathocrinus swallovi – both have columns indicating current that buried them came from the left in this photo.

Cyathocrinites multibrachiatus (Lyon & Casseday) – New Providence Shale, Middle Mississippian, KY – C,P,1

Cyathocrinites multibrachiatus body plates
Cyathocrinites multibrachiatus body plates

Cymbiocrinus sp.# – Bangor Fm., Chesterian, Upper Mississippian, Colbert Co., Alabama – C,2

Cymbiocrinus sp. in limestone
Cymbiocrinus sp. in limestone

Column –geodized                              Middle Devonian                            IN                           S,1

Dichocrinus simplex Shumard, Middle Mississippian, IN,KY – C,1,2

Dichocrinus simplex  (left) and Synbathocrinus swallovi (right) - both from the Muldraugh Fm. of Hardin Co., Kentucky
Dichocrinus simplex (left) and Synbathocrinus swallovi (right) – both from the Muldraugh Fm. of Hardin Co., Kentucky

Dizygocrinus montgomeryensis (Worthen)# – Muldraugh Formation, Middle Mississippian, Hardin Co., KY – C,1,2

Dizygocrinus montgomeryensis in limestone
Dizygocrinus montgomeryensis (Worthen)

Dizygocrinus sp. – Muldraugh Formation, Middle Mississippian, Hardin Co., KY – C,1,2

Dolatocrinus bellulus - basal cup
Dolatocrinus bellulus – basal cup
Dizygocrinus sp. calyx (compressed, ventral view)
Dizygocrinus sp. calyx (compressed, ventral view)

Dolatocrinus bellulus (Miller & Gurley)# – Beechwood Ls., Givetian, Middle Devonian, Clark Co., IN & Jefferson Co., KY – S,1

Dolatocrinus bulbaceous MIller & Gurley# – Beechwood Ls., Givetian, Middle Devonian, Clark Co., IN – S,1

 Dolatocrinus bulbaceous - from the deeply weathered Beechwood Limestone
Dolatocrinus bulbaceous – from the deeply weathered Beechwood Limestone

Dolatocrinus grandis Miller & Gurley# – Beechwood Ls., Givetian, Middle Devonian, Jefferson Co., KY – S,2

Dolatocrinus grandis Miller & Gurley
Dolatocrinus grandis Miller & Gurley

Dolatocrinus stem / columnals – Beechwood Limestone, Givetian, Middle Devonian, IN,KY – C,S,1,2

Dolatocrinus columnals
Dolatocrinus columnals

Dolatocrinus vetustus – one of the most beautiful species, Beechwood Limestone, Givetian, Middle Devonian, Clark Co., IN – S,1,2

Dolatocrinus vetustus has highly ornate plates - Mark Schneider photo
Dolatocrinus vetustus – Mark Schneider photo

Ectenocrinus simplex (Hall) – Kope Fm., Upper Ordovician, KY – C,1,2

Ectenocrinus simplex – possibly the most common crinoid of the Tri-state Ordovician

Eucalyptocrinites crassus (Hall) – is the most common crinoid in the Waldron Shale, Middle Silurian, Clark Co., IN – C,1,2

Eucalyptocrinites crassus with arm struts

Eucalyptocrinites elrodi (Miller) = E. tuberculatus (Miller & Dyer) – has ornamented body plates, Waldron Shale, Middle Silurian, Clark Co., IN – C,1

Eucalyptocrinites elrodi – with arm struts preserved but no arms

Eucladocrinus (possibly E. millebrachiatus) stem (oval disks) – form is similar to Platycrinites, but their columnals aren’t as wide. Middle Mississippian, Hardin Co., KY – C,1

Eucladocrinus column – large oval columnals

Eupachycrinus sp.# – Indian Springs Shale, Chesterian, Upper Mississippian, Crawford Co., Indiana – C,1,2

Eupachycrinus sp. – this photo doesn’t show diagnostic details

Fifeocrinus popensis (Worthen)# – Beech Creek Fm., Chester, Upper Mississippian, Breckinridge Co., Kentucky – C,2

Fifeocrinus popensis – fine calyx

Forbesiocrinus sp.# – is a flexible crinoid that isn’t as common as Taxocrinus, from the Middle Mississippian, IN – C,2

Gaulocrinus trautscholdi body plates
Forbesiocrinus sp – a 15 mm tall calyx

Gaulocrinus trautscholdi  (Wachsmuth & Springer) – New Providence Shale, Middle Mississippian, KY – C,1 & P,1

Halysiocrinus tunicatus (Hall) – New Providence Shale, Middle Mississippian, KY – C,1;

Halysiocrinus tunicatus body plates

Holdfast – Eucalyptocrinites – radiate out in or on a substrate, Waldron Shale, Middle Silurian, Clark Co., IN – C,2

Holdfast of Eucalyptocrinites on the cup of a E. crassus

Holdfast – Gilbertsocrinus# – look like coiled rope, from the Devonian and Mississippian, IN, KY – C,S,1,2

Gilbertsocrinus holdfast – silicified
Lichenocrinus – a group of holdfasts on Rafinesquina brachiopod

Holdfast – Lichenocrinus sp. – plated, belongs to Cincinnaticrinus, a genus that were generally small. Upper Ordovician, KY – C,2

Holdfasts, multi-cirri – Beech River Formation, Middle Silurian  Decatur Co., Tennessee – C,1

Collection of multi-cirri holdfasts

Holdfasts, multi-cirri – had columns radiating out into the sediment to anchor the animal, Jeffersonville & Beechwood Limestones, Middle Devonian, Clark Co., IN – S,1

A multi-cirri crinoid holdfast

Holdfasts, multi-cirri – had columns radiating out into the sediment to anchor the animal, New Providence Shale, Middle Mississippian, Jefferson Co., KY – C,S,1,2  

A multi-cirri crinoid holdfast

Holdfast, Podolithus – is a crinoid holdfast form genus described by Sardeson in 1908. Discoidal with a depressed center, it’s common to many genera over time and is often preserved attached to a substrate. In the Beechwood Limestone, they are often weathered loose.It is found in the Ordovician, Silurian and Devonian. IN, KY – C,S,1,2

Podolithus attached to Favosites placentus in the Beechwood Ls.
Podolithus from the Middle Devonian Beechwood Ls., Clark Co., IN
Podolithus on a bryozoan from the Kope Fm., Upper Ordovician, Carroll Co., KY. Holdfast for Anomalocrinus. FOV ~2cm

Hypselocrinus sp.# – Indian Springs Shale, Chesterian, Upper Mississippian, Crawford Co., Indiana – C,1,2

Hypselocrinus sp. – cup, FOV ~2 cm

Hyrtanocrinus sp.# – Paoli Ls., basal Chesterian, Upper Mississippian, from the Talarocrinus patei beds near Sample, Breckinridge Co., KY – C1,2

Hyrtanocrinus sp. with plates like fingerprints

Iocrinus sp.# – widespread in the Upper Ordovician of the Tri-state area, IN-KY-OH C,1,2

Iocrinus with a star-shaped base

Lampterocrinus tennessensis (Roemer)# – camerate crinoid from the Beech River Fm., Middle Silurian, Decatur Co., Tennessee. C,1

Lampterocrinus tennessensis has plates that look like a cystoid, but it’s a crinoid.

Lecanocrinus pisiformis (Roemer) – is a small crinoid found in the glades of the Beech River Fm., Middle Silurian, Decatur Co., TN – C,1; Rarer than Pisocrinus.

Lecanocrinus pisiformis – is a small species

Lecanocrinus pusillus Hall# – from the Waldron Shale, Middle Silurian, it’s larger than the Beech River species but is still small. Clark Co., IN – C,1,2

Lecanocrinus pusillus – this is not a great photo. Cup is ~1 cm wide

Lecocrinus? springeri Kammer# – New Providence Shale, Middle Mississippian, KY – C,1

Lecocrinus? springeri body plates

Lyriocrinus melissa (Hall) – is more likely to be found with arms in the Waldron Shale, Middle Silurian,   than any other species. Clark Co., IN – C,1,2

Lyriocrinus melissa with partially preserved biserial arms

Melocrinus obconicus Hall – Waldron Shale & Louisville Ls., Middle Silurian, IN – C,2

Melocrinus obconicus – prepped.

Metichthyocrinus tiaraeformis# – has a beautiful basal cup, New Providence Shale, Middle Mississippian, Jefferson Co., KY – C,1

Metichthyocrinus tiaraeformis basal cup – 3 cm FOV

Myelodactylis cf. M. ammonis Bather – Waldron Shale, Middle Silurian, IN – C,2

Myelodactylis cf. M ammonis – unprepped
Myelodactylis cf. M ammonis – unprepped
Myelodactylis cf. M ammonis – prepped – tiny calyx is in the center

Myelodactylus convolutus Hall – consisted of a knotted stem that used another crinoid stem as an anchor. Waldron Shale, Middle Silurian, IN – C,2

Myelodactylus convolutus – prepped by Chris Wright
Myelodactylus convolutus

Paradichocrinus ramus Ausich, Goldstein & Yates# – a second new species from our research published in 2000 on the crinoids from the Muldraugh Fm., Middle Mississippian, Hardin Co., Kentucky. C,2

Paradichocrinus sp. through a microscope, field <3cm vertical
Paradichocrinus ramus Ausich, Goldstein & Yates# - a second new species from our research published in 2000
Paradichocrinus ramus Ausich, Goldstein & Yates# – a second new species from our research published in 2000

Paradichocrinus sp.# – from the Muldraugh Fm., Middle Mississippian, Hardin Co., Kentucky. C,2

Pentaramicrinus sp.# – Upper Mississippian, Colbert Co., Alabama – C,1,2

Pentaramicrinus sp. = partial calyx
Pentaramicrinus sp. – basal cup, FOV ~2.5 cm

Periecocrinus christyi (Hall) – Waldron Shale, Middle Silurian, Clark Co. Indiana, C,1,2  

Periecocrinus christyi – dampened to show plates better

Phanocrinus alexandrii Strimple# – Upper Mississippian, IN, KY – C,1

Phanocrinus alexandrii – cup and base of the arms

Phanocrinus formosus (Worthen)# – Upper Mississippian, IN, KY – C,1

Phanocrinus formosus – cup and base of the arms

Phanocrinus parviramus Sutton & Winkler# – Indian Springs Shale Mbr., Big Clifty Fm., Upper Mississippian, Crawford Co., IN – C,2

Phanocrinus parviramus

Phanocrinus sp. – Upper Mississippian, IN,KY – C,1,2

Phanocrinus sp. (basal cup) 

Phanocrinus wetherbyi # – Indian Springs Shale Mbr., Big Clifty Fm., Upper Mississippian, Crawford Co., IN – C,2

Phanocrinus wetherbyi – a large specimen

Pisocrinus quinquelobus Bather – basal cups are typical, Beech River Fm., Middle Silurian, Decatur Co., TN – C,1,2

Pisocrinus quinquelobus -basal cup

Platycrinites bonoensis (White) – cups are common in the Salem Limestone, Middle Mississippian, Hardin Co., KY – C,1,2

Platycrinites bonoensis

Platycrinites hemisphaericus (Meek & Worthen)# – possible body plates from the New Providence Shale, Middle Mississippian, KY – C,1

Platycrinites georgii # – Muldraugh Fm., Middle Mississippian, Hardin Co., KY – C,1,2 (See photo with Cosmetocrinus gracilis, above)

Platycrinites hemisphaericus? body plates

Platycrinites huntsvillae # – Muldraugh Fm., Middle Mississippian, Hardin Co., KY – C,1,2

Platycrinites huntsvillae – a complete crown

Platycrinites penicillus # – columnals, Middle – Upper Mississippian, Hardin Co., Illinois – C,S,1,2

Platycrinites penicillus – distinctive spiny, oval columnals

Platycrinites planus (Owen & Shumard)# – body plates – New Providence Shale, Middle Mississippian, KY – C,1

Platycrinites planus, etc., body plates

Platycrinites sp. – oval columnals,  New Providence Shale, Middle Mississippian, KY – C,1,2

Platycrinites sp. columnals

Platycrinites saffordi (Hall)# – New Providence Shale, Middle Mississippian, KY – C,1

Platycrinites saffordi – ornamented basal plate

Pterotocrinus acutus Wetherby – wing plates,  Upper  Mississippian, Crawford Co., IN, Grayson Co., KY – C,1

Pterotocrinus acutus wing plates

Pterotocrinus acutus variant 

Pterotocrinus acutus variant 

Pterotocrinus acutus Wetherby# – whole body, Glen Dean Fm., Upper Missisippian, Grayson Co., Kentucky

Pterotocrinus acutus has five elongate wing plates and usually has a coprophagous Platyceras snail at the anal tube, as seen in this specimen.
Pterotocrinus acutus basal view

Pterotocrinus bifurcatus variant of P. acutus   

Wing plate of Pterotocrinus bifurcatus variant of P. acutus

Pterotocrinus depressus Lyon & Casseday – wing plates are common, Upper Mississippian, Crawford Co., IN, Grayson Co., KY – C,1,2

Pterotocrinus depressus wing plates

Pterotocrinus spatulatus – another acutus variant? Upper Mississippian, Crawford Co., IN, Grayson Co., KY – C,1,2

Pterotocrinus spatulatus wing plate

Pterotocrinus vannus Sutton – wing plates are common, Upper Mississippian, Crawford Co., IN, Grayson Co., KY – C,1,2         

Pterotocrinus vannus

Scytalocrinus robustus (Hall) – Muldraugh Fm., Middle Mississippian, Hardin Co., KY – C,2

Scytalocrinus robustus – finding this specimen led to the discovery of a “crinoid quarry” in Hardin Co., KY

Springerocrinus sp.# – body plates, Middle Mississippian, KY – C,2

Springerocrinus sp., body plates – 3 cm view

Strimplecrinus coxanus (Worthen)# – is a very small species with ornamented plates, Salem Ls., Middle Mississippian, IN, KY – C,1,2

Strimplecrinus coxanus – ornamented basal cup, FOV ~ 1 cm

Synbathocrinus sp. (probably swallovi) body plates – Middle Mississippian, KY – C,1

Synbathocrinus sp. (probably swallovi) body plates

Synbathocrinus swallovi Hall – has simple straight arms and a small cup, Middle Mississippian, KY – C,1,2

Synbathocrinus swallovi basal cup, silicified, FOV ~2.5 cm

Talarocrinus patei (Miller & Gurley) – Paoli Ls., basal Chesterian, Upper Mississippian, Breckinridge Co., KY – C,1,2

Talarocrinus patei calyx with column from a mass mortality

Talarocrinus simplex (Shumard)# – Salem Ls., Middle Mississippian, IN, KY – C,1,2

Talarocrinus simplex – is mostly found as a basal cup, FOV ~2.5 cm

Taxocrinus colletti White – is the most common species of flexible crinoid in the Middle Mississippian Borden Delta sediments. IN, KY – C,1,2

Taxocrinus colletti White – view of arms, a large specimen prepared by Chris Wright.
Taxocrinus colletti – base of crown with a short column segment attached.

Taxocrinus whitfieldi (Hall) – a common though usually fragmented species, widespread in the Upper Mississippian of eastern N. America. Crawford Co., IN – C,1,2

Taxocrinus whitfieldi – wet to show detail, FOV ~2.5 cm

Tholocrinus spinosus Wood – one of several genera of spiny crinoids in the Upper Mississippian, IN, KY – C1,2

Tholocrinus spinosus from Grayson Co., KY collected by Mark Palatas.

Zeacrinites wortheni Hall# – Indian Springs Shale Mbr., Big Clifty Fm., Upper Mississippian, Crawford Co., IN – C,2

Zeacrinites wortheni – Genus Zeacrinites was named by Gerard Troost because his calyx resembled an ear of corn (Zea)

Stems                                                     Mid-Upper Mississippian                  KY                         C,S, Fluorite replaced/encrusted